Once water crosses from xylem to phloem, it flows through the trunk toward the energy consuming parts of the tree. The water becomes solvent for sucrose, and the flowing material in phloem is sap. The phloem is essentially one long pipe of constant radius, with length equal to the height of the tree.
Compared to water, sap is significantly more viscous and thus will exert friction on itself that slows its flow. As with the membrane, we can find a relation between osmotic pressure and volume flow that we call the hydraulic resistance of the sap in the stem, .
We can calculate this resistance exactly but it is a little involved, so we'll first try to derive its functional form by scaling arguments. As the great John Wheeler once said, "Never make a calculation until you know the answer."
We expect the area of the cross section, the length of the pipe , the pressure across the pipe , and the viscosity of the liquid to come into play.
Use physical intuition and dimensional analysis to come up with a relation, between flow rate , viscosity , pipe length , cross sectional area , and pressure . Use limiting cases to check your hypothesis.
Details and Assumptions:
This section requires Javascript.
You are seeing this because something didn't load right. We suggest you, (a) try
refreshing the page, (b) enabling javascript if it is disabled on your browser and,
finally, (c)
loading the
non-javascript version of this page
. We're sorry about the hassle.
The flow rate u is directly proportional to cross-sectional area A and the force due to pressure Δ p A ; and inversely proportional to viscosity η and tree height h . Therefore the expression is: u = η h A 2 Δ p